1964] TURNER & ItORNE : MACIIAERANTItERA SECT�9 PSILACTIS 331 �9 1962a. Interspecific hybridization in Haplopappus and its bearing on chromosome evolution in the Blepharodon section. Am. Jour. Bot. 4:9: 119-132. �9 1962b. Intergeneric hybridization between Psilactis and Machaeranthera, Am. Jour. Bot. 49: 676. (Abstract). Raven, P. It., Solbrig, O. T., Kyhos, D. W., & Snow, R. 1960. Chromosome numbers in Compositae. I. Astereae. Am. Jour. Bot. 47: 124-132. 8hinners, L. tI. 1950. Notes on Texas Compositae V. Field Lab 18: 32-42. Turner, I3. L, Beaman, ~. It., & Rock, I-I. F. L. 1961. Chromosome numbers in the Com- positae V. Mexican and Guatemalan species. Rhodora 63: 121-129. , tlllison, W. L., & King, 1~. 1~. 1961. Chromosome numbers in the Compositae. IV. North American species, with phyletie interpretations. Am. Jour. Bot. 48: 216-223. , & Johnston, lYI. (J. 1961. Chromosome numbers in Compositae I I I . Certain Mexican species. Brittonia 13" 64-69. , Powell, M., & King, R. M. 1962. Chromosome numbers in the Compositae VI. Additional Mexican and Guatemalan species�9 Rhodora 64: 251-271. A NEW ANTIDAPI-INE FROM ECUADOR JOB KUIJT University of British Columbia, Vancouver. In comparison with other loranthaceous genera, Antidaphne has led a rather undisturbed existence in botanical l i terature since its description in 1838 by Poeppig and Endlicher. I t was not unti l the end of the century that a second species (v. Tieghem 1895) joined the original A. viscoidea. Van Tieghem erected the genus Stachyphyllum upon the second species, which subsequently was in- corporated in Antidaphne as A. fendleri (v. Tiegh.) Engler (1897). More than half a century later Rizzini (1956) attempted a brief survey in which he added two more species. The above summarizes the entire botanical history of the genus, with the exception of occasional appearances of A. viscoidea in floristic works. The present paper adds one more species, and undoubtedly its most distinct one�9 Ant idaphne orbicularis Kui jt , sp. nov. P lanta robusta, caulibus teretibus vel leviter eompressis, foliis alternantibus �9 Fol ia orbieularia, frondosa diametro usque ad 18 em, basi eordata, petiolo bre- vissimo robustissimo, venatione palmata. Spiea squamis deeiduis, statu frueti fero fere elongata (usque ad 4 em in spiels determinatis), saepe exerescens, spieas alias et ramulos vegetativos gerens. Fol ia apiealia spicarum diametro ad 4 cm, spiearum exereseentium multo majora. Flores feminei ca 1 mm longi; f ructus baceatus, ovatus, monospermus, diametro 4 )< 3 mm; semen diametro 3 )( 2 mm, ellipticum, faseieulis simplicibus a basi ad apieem percurrentibus. A large plant with str ikingly large (up to 18 em in diameter) orbicular foliage leaves with cordate base and very short, stout, f lattened petiole giving rise to about 5 major veins in a palmate fashion ; phyl lotaxy alternate �9 Branches often terete, but spikes and vigorous branches compressed or even keeled, inter- nodes then about 5 cm long�9 Spikes with deciduous scales, the former axil lary in groups of hal f a dozen or more�9 Axi l lary region of large foliage leaves a cushion of 30-40 buds representing spikes to be developed in successive genera- tions; one or two vegetative shoots may take their origin f rom the center of such a cushion. Pistil late spikes expanding in fruit, when determinate to about 4 cm, when becoming percurrent bearing fruit only near the base, producing BRITTONIA 16: 331--333. Jul 1964. 332 BRITTONIA [VOL. 16 Fla. 1-2. Holotype of Antidaphne orbicularis. FIG. 1. Habit. Length of label 11.5 em. FIG. 2. Same, detail of inflorescences. more spikes and other branches higher up. Pistillate spikes with up to a dozen fertile, eaducous bracts, the axi! of each of which bearing 3-5 fl~owers in a transverse row; central flower or fruit largest, the lateral ones successively smaller. The apex of a young spike bears 1-3 minute bract-like organs which expand into small (up to 4 cm in diameter) foliage leaves, of the same shape and venation as large ones, when the spike elongates. Pistillate flower about 1964] KUI JT : :NEW ECUADOREAN ANTIDAPttNE 333 1 mnl long; style prominent even in fruit, stigma small, eapitate. Fruit an ovate, one-seeded berry 4 X 3 mm; seed 3 }< 2 mm, elliptical, with about 6 unbranched vascular bundles connecting both poles. TYPE: C. H. Dodson and L. B. Thien 2123, Ecuador, Prov. Comar, km 110 from Duran; tropical cloud forest; elev. 1300 m. Parasite on tree. Holotype, MO. Isotype, UBC. The type collection is a pistillate plant, and one can only guess what the staminate flowers may be like. Floral morphology is exceedingly constant within each genus of the Viscoideae, fortunately; so that we can be reasonably certain that the following description of the staminate flowers of A. viscoidea from Costa R~ca is close to that of A. orbicularis. In the former, the staminate spike is slightly longer than the pistillate one; it also has caducous bracts and about the same number and arrangement of flowers. Each staminate flower is reduced to 2-4 stamens around a central cushion, the entire unit raised on a pedicel of about the length of a stamen. In some Andean material (as in the Bang collec. tion mentioned below) the number of stamens per flower may be as many as eight. As in the pistillate spike, several flowers may be developed per axil. The present species also fills the major gap in the geographic distribution of the genus. Antidaphne has been reported from Peru by Poeppig and Endlicher (1838) as well as Maebride (1937); from Brazil, Venezuela, and Bolivia by Rizzini (1956) ; from Panama by Rizzini (1960) ; from Costa Rica by Trelease (1937) and subsequently by Kuijt (1964); and from Guatemala by Standley and Steyermark (1946). Colombia is represented by a specimen, apparently A. viscoidea, at the Gray Herbarium (Bang 2257), and Mexico by Matuda 3997 (F). A. orbic~daris is the first record of the genus in Ecuador. Again extrapolating from Costa Rican observations on A. viscoidea, the present species is likely to develop epicortical roots in youth, these being gradu- ally replaced by a greatly enlarged primary attachment. The proliferation of spikes into vegetative shoots described here for A. orbicularis, is also clearly visible in the isotype of A. fendleri at the Gray Herbarium (Fendler 1125), but has not been encountered elsewhere. Indeed, Oliver described the situation quite accurately in his important morphological remarks on the genus (1864). A. orbicul~ris differs strikingly from all other described material in its large, orbicular leaves with palmate venation. I am obliged to Dr. Karel U. Kramer for the Latin diagnosis. L ITERATURE C ITED F.ngler, A. 1897. Loranthaceae . ~rn Eng ler & Prant l , Natfir l . B f lanzenfam. , Naehtr . 1 :138 Kuijt, Y. 1964. A revision of the Loranthaceae of Costa Rica. Pot. Jahrb. (in press). l~Iacbride, J. F. 1937. Flora of Peru (Loranthaceae). Publ. Field :~s Pot. 13: 375-416. Oliver, D. 1864. Notes on the Loranthaceae, with a synopsis of the genera. Jour. Linn. Soe Pot. 7: 90-106. Poeppig, E., and Endlicher, S. 1838. Nova Gen. et Spec. 2: 70, t. 199. Rizzini, C. T. 1956. Pars specialis prodromi monographiae Loranthaeerum Brasiliae terr~ rumque finitimarum. Rodrigu6sia 18-19 (30-31): 87-234. �9 1960. Loranthaceae. In Woodson, R. E., Jr., and Schery, R. W., Flora of Panama. Ann. :Missouri. Pot. Gard. 47: 263-290. Standley, P. C., and Steyermark, J. A. 1946. Flora of Guatemala. Loranthaceae. Fieldiana Pot. 24: 62-86. Tieghem, P. van. 1895. Sur les genres Ba~icarpus, Stachyphyllum et Antidaphne de la sous- famille des Viscoid~es. Bull. Soc. Pot. Fr. 42: 562-573. Trelease, W. 1937. Loranthaceae. In Standley, P. C., Flora of Costa Rica. Publ. Field :Mus. Pot. 18: 402-408.